Isalo National Park. East side. Eastern edge of park at 34 km from Ranohira on Route Nationale 7, W of road to Beroroha, near Kelivozana., Ihorombe, Madagascar, Africa
Sava Region. Reserve Naturelle de Marojejy; along the Manantenina River, N of the 2nd camp on the trail to the summit of Marojejy Est., Sava, Madagascar, Africa
Fivondronana Fianarantsoa, Ranomafana National Park, Talatakely area, going up along Namorona River, on the intersection between Sakaroa stream and Namorona River., Vatovavy Fitovinany, Madagascar, Africa
Between Ambositra and Ambatofinadrahana. On Route Nationale 35, 12 km W of Ivato. Uapaca bojeri woodland, above waterfall., Amoron'i Mania, Madagascar, Africa