Brown, R. M., Guttman, S. I. 2001. Phylogenetic systematics of the Rana signata complex of Philippine and Bornean stream frogs: reconsideration of Huxley's modification of Wallace's Line at the Oriental-Australian faunal zone interface. 76 427.
Type Status:
Paratype
Place:
Puerto Galera Municipality, Barangay Villaflor, ca. KM 15 (from Puerto Galera) on Calapan - Puerto Galera Road, Tamaraw Waterfalls, Oriental Mindoro, Mindoro Island, Philippines, Asia