Heyer, W. R., Wolf, A. J. 1989. Physalaemus crombiei (Amphibia: Leptodactylidae), a new frog species from Espirito Santo, Brazil with comments on the P. signifer group. 102 (2): 500.
Type Status:
Paratype
Place:
Santa Teresa, near, edge of Reserva Biologica Nova Lombardia, Espirito Santo, Brazil, South America