Heyer, W. R., Garcia-Lopez, J. M., et al. 1996. Advertisement call variation in the Leptodactylus mystaceus species complex (Amphibia: Leptodactylidae) with a description of a new sibling species. 17 25, Figure 13.
Type Status:
Paratype
Place:
Puerto Maldonado, ca. 30 km (airline) SSW of, Tambopata Reserve, Explorer's Inn, Madre de Dios, Peru, South America