Lachner, E. A. & Jenkins, R. E. 17 Aug 1971 ["1971"]. Smithsonian Contributions to Zoology. No. 85: 17.
Type Status:
Paratype
Place:
Virginia, Blackwater River of Roanoke Drainage at Route 22 bridge about 4 air miles northeast of Rocky Mount, Franklin County., Franklin County, Virginia, United States, North America