Molina, J. E., Struwe, L. 2008. Syst. Biodivers. 6 (4): 490.
Type Status:
Isotype
Place:
Zamora-Chinchipe: Cordillera del Condor, 18.4 km E of Los Encuentros (as measured from the bridge crossing the Rio Nangaritza E of the city center) in direction of the Paquisha Alta military camp., Zamora-Chinchipe, Ecuador, South America - Neotropics