Leonard, E. C. & Smith, L. B. 1964. Rhodora. 66: 332.
near Arroyo Bravo on road to Divisoria, near Tingo María, San Martín, Peru, South America - Neotropics
16 Dec 1949
"Sheet I", this sheet annotated as holotype by Leonard. "Sheet 2", USNH 2026628, was annotated by Leonard as "Isotype" but is to be considered as second sheet of holotype (fide D. Koenemann, based on discussion with J. McNeill, email comm. June 2011).